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Cheesemonger's Mac and Cheese

['1 1/2 cups coarsely grated Gruyère cheese', '1 1/2 cups coarsely grated sharp cheddar cheese (about 6 ounces)', '1 1/2 cups diced rindless Brie (cut from 1-pound wedge)', '5 tablespoons butter, divided', '1/4 cup all purpose flour', '2 teaspoons chopped fresh thyme leaves', '3/4 teaspoon (scant) nutmeg', '4 cups whole milk', '1 3/4 cups fresh breadcrumbs made from crustless French bread', '1 pound penne pasta', '8 teaspoons whipping cream (if making 1 day ahead)']

Mix all cheeses. Set aside 1 cup for topping; cover and chill. Melt 4 tablespoons butter in large saucepan over medium heat. Add flour and stir until mixture turns golden brown, about 4 minutes. Add thyme and nutmeg. Gradually whisk in milk. Simmer until thickened and smooth, stirring often, about 4 minutes. Add cheeses from large bowl. Stir until melted and smooth.
Melt 1 tablespoon butter in heavy large skillet over medium-high heat. Add breadcrumbs; toss. Stir until golden, about 2 minutes. Transfer to plate.
Preheat oven to 375°F. Cook pasta in boiling salted water until tender but firm to bite. Drain. Transfer to large bowl. Pour cheese sauce over; toss. Divide among eight 1 1/4-cup custard cups. Sprinkle with 1 cup cheese. Place cups on rimmed baking sheet. DO AHEAD: Can be made 1 day ahead. Cover with foil; chill. Drizzle each with 1 teaspoon cream. Cover with foil; bake 15 minutes. Uncover. Sprinkle partially baked chilled or just assembled cups with breadcrumbs. Bake pasta until beginning to bubble and tops are golden, about 20 minutes.

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